Update guys fuel system bled this evening.
No fault codes From diagnostics.
Relays below seat checked and working.
Tomorrow I’m going to bypass the immobiliser on the instrument cluster and see if that does the job.
 
Hope these attachments help. (but are taken from a manual for left hand drive). Looking at the page for Principles of Operation, the starter relay is energised if the vehicle gets a valid mobilised signal. As your starter motor is working, I would say the vehicle is not immobilised and you should be looking elsewhere for the problem.
 

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Thanks wimblowdriver.
I’ve decided to leave it a few days it’s doing my head in. Went out this morning battery totally drained like my love for defenders at present. Went out and bought a noncom booster which aren’t cheap.
It might surprise me and turn over a new leaf for new year. Well I’m hoping so.
 
just because the battery is new does not mean it's any good check the voltage before condemning a list of item you might think it is IF THE ALARM UNIT WAS ACTIVE YOU WOULD HAVE NO CRANK AND NO FUEL DELIVERY
ANTI THEFT Defender RHD VIN 732615 to 751062-75 2007.png
 
Been thinking about the OP's problem, which started after a refuel. Something is inhibiting starting, but not the immobiliser as the engine cranks over. I have the handbook for the 2.2tdci and there is a paragraph in there regarding running out of fuel (assuming the 2.4 has the same characteristics). If the fuel level falls below a minimum level the engine runs at reduced power, followed by the engine stopping. After refuelling it has to be restarted with the following procedure;
1. With the brake pedal depressed, turn and hold the engine start key and crank the engine for 5 seconds.
2. Release the key.
3. With the brake pedal still depressed, turn the key and crank the engine. The engine should start within approximately 5 seconds.
If the engine does not start, pause for 10 seconds and repeat. Do not crank for more than 30 seconds or damage to the fuel pump may occur.
A long shot but worth a try?

Another item to look at is the Overide Inertia Switch located on the bulkhead, which might have been activated and may need resetting, or its fuse might need replacing. (This cuts off the fuel supply (via the ECU) in the event of a collision). Again, a long shot but worth checking.
 
Hi guys happy new year to you all.
I will look for this inertia switch where is it located exactly. I’ve tried to bypass immobiliser no joy. Yes correct fuel. The way it stands now is since the battery drain. We have ignition lights and it cranks for a second then nothing. The battery has been boosted and sitting at 12.49 volts.
I think since the battery drain the immobiliser as kicked in but I also now think I have two problems the immobiliser and a fuel starvation issue.
 
If it cranks, dash lights work etc it is NOT immobilised. I suggested previously, you might need to resynchronise the key fob after a vehicle battery failure.
 
Quick up date for you guys that have been so helpful.
The glow plug relay had jammed on which we discovered is draining the battery and a couple of glow plugs aren’t working.
The engine is turning over but not firing up. Fuel rail has pressure but injectors aren’t injecting conclusion is the car is immobilised and has a faulty AS10 which isn’t allowing ECU to be reset. So the ECU is being flashed to remove the immobiliser and new alarm immobiliser will be installed once we get her running. Slowly moving forward hopefully in the correct direction.
 

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