Not 100% about that. What I do know is that Disco 2 wheels don't fit on a Ninety.
I know this because at one time I had both a Ninety and a Disco 2, and out of curiosity, I tried a wheel off the Disco on the Ninety.
Not sure how that relates to late model Defenders, as I have never had one.
Stud patterns stopped the same on the LR but changed on the disco/RR

I was amazed the “new”defender rims fit on D3/D4
 
I know everyone here is talking about whether the wheels will physically fit the hubs but you also need to look at the weight rating of the wheels. Less important if you have a 90 but if you have a 110 or a 130 then there are plenty of the wheels are not rated to the same as the axel weights of the larger vehicles. From memory I think only boost alloys are rated for a 110, and freestyle alloys definitely aren't.
 

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