Disco definetly wins it on the comfort front, off-road my 'standard' disco grounds out on the tow bar a bit, but, it has to be kept 'standard' to keep the gf happy. It does, however, make an excellent tow vehicle for taking the series long distances.
The series i find is actually a lot more fun than a defender (yes i have owned one), not as easy to extremly modifiy it, but, bouncing around on leafs in something which is nearly 50 years old and watching people in 5 year old defenders blowing up bits of engines and getting stuck in things you can drive through / up is extremly satisfying.
comes down to what you need, what you can afford and what floats your boat.